BMP2, CAC, CLDN2, CLDN4, CLDN7, CLDN8, Claudin 2, Claudin 7, Claudin 8, Claudin protein, Cldn2, Cldn3, Cldn4, Cldn7, Cldn8, Colon epithelial cell, FITC, Human, IBD, In vitro colon organoid culture, Mice, Mouse, RNA, SMAD4, SMAD4+, Smad4, Smad4 gene, TGF, TGF family, TGF1, colitis-associated cancer, colon, colon cancer, colon epithelial cells, gene, healthy controls, human, inflammatory bowel disease, mRNA, mice, mouse, nascent RNA, polarized SMAD4+ colonoids, tight